41. Geranium krameri Franchet & Savatier, Enum. Pl. Jap. 2: 306. 1878.
突节老鹳草 tu jie lao guan cao
Geranium sieboldii Maximowicz.
Perennials. Rootstock ± vertical, 5-12 mm in diam., not tuberculate, with a fascicle of long thickened roots. Stem (25-)50-129 cm tall, erect, not rooting at nodes, with 0.4-0.8 mm retrorse appressed or patent nonglandular trichomes. Stipules lanceolate, distinct or connate at least at base. Leaves opposite; petiole with 0.3-0.8 mm retrorse appressed or patent nonglandular trichomes; leaf blade 5.6-11.9 cm, palmately cleft, ratio of main sinus/middle segment length = (0.81-)0.83-0.87, pilose with appressed nonglandular trichomes; segments 5 or 7, rhombic, 4.3-8.8 mm wide at base, 5-14-lobed in distal half, ratio of second sinus/middle segment length = 0.18-0.32. Cymules solitary, 2-flowered; peduncle 1.2-8.8 cm. Pedicel 2.3-5.5 cm, with 0.1-0.7 mm retrorse appressed or patent nonglandular trichomes; bracteoles lanceolate. Sepals (6.2-)7-10 mm, mucro 0.6-1.3 mm, ratio of mucro/sepal length = 0.1-0.15, outside with 0.3-0.7 mm antrorse ± appressed nonglandular trichomes, inside glabrous. Petals light pink, (1-)1.1-1.4(-1.5) cm, erect to patent, outside basally with trichomes, inside basal 1/3-1/2 with trichomes, margin basally ciliate, apex rounded. Staminal filaments pinkish, lanceolate, base slightly dilated, abaxially pilose and proximal half ciliate, trichomes 0.2-0.7 mm; anthers brown, 1.6-2.1 mm. Nectaries 5, hemispheric, abaxially glabrous, apex with a tuft of trichomes. Stigma brown. Fruit 2.6-3.1 cm, erect when immature; mericarps smooth, with a basal callus, with 0.1-0.4 mm ± patent nonglandular trichomes and apically usually with 0.4-0.7 mm patent glandular trichomes; rostrum 1.6-2 cm, with a 1.6-2.6 mm narrowed apex; stigmatic remains 4.6-6.6 mm. Seeds 1.6-2.2 mm. Fl. Jul-Aug, fr. Aug-Sep. 2n = 28.
Scrub, meadows, weedy areas in farmlands; 600-1200 m. Heilongjiang, Jilin, Liaoning [Japan, Korea, Russia (Far East)].
The indumentum of this species is variable, with patent trichomes in some specimens and retrorse and appressed in others. As in Geranium dahuricum and G. soboliferum, trichomes of petals are ± dense on the base of the inside and extending upward on veins. Geranium yesoense Franchet & Savatier is a related species endemic to Japan, Sakhalin, and Kurile Islands, with nectaries glabrous, longer petals, similar leaf blade divisions, and rootstock with thickened and not fasciculate roots.