Non-Migrant: No. All populations of this species make significant seasonal migrations.
Locally Migrant: Yes. At least some populations of this species make local extended movements (generally less than 200 km) at particular times of the year (e.g., to breeding or wintering grounds, to hibernation sites).
Locally Migrant: Yes. At least some populations of this species make annual migrations of over 200 km.
Landlocked populations may migrate up to about 50 miles upstream for spawning, anadromous populations up to 200 miles (Scott and Crossman 1973).